The agency did not respond to requests for comment on its record monitoring animal research experiments nationally.
The USDA inspector general has published at least three reports since 2014 critical of the agency’s lax oversight, though its criticism dates back to the 1990s.
Neuralink says on its website that it champions animal welfare and tries to reduce animal testing where possible.
Two academic studies conducted in 2009 and 2012 found that animal research committees approved between 98% and 99% of experiments proposed by researchers.
Envigo was made to sign the consent decree giving up the beagles only after the USDA inspector general and the Justice Department investigated and found evidence of inhumane treatment.